What is this consistent symbol in the Spanish coat of arms?

What does the symbol on the Spanish coat of arms mean?

:: Meaning of the Spanish Coat of Arms ::



The four quadrants signify the four kingdoms which came together to form a unified Spain in the late 1400’s. Castile represented by a castle, León represented by a lion, Aragon represented by vertical red and yellow stripes, and Navarre represented by linked chains.

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